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Who discovered chromosomes?

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Visual discovery of chromosomes. Textbooks have often said that chromosomes were first observed in plant cells by a Swiss botanist named Karl Wilhelm von Nägeli in 1842. However, this opinion has been challenged, perhaps decisively, by Henry Harris, who has freshly reviewed the primary literature. In his opinion the claim of Nägeli to have seen spore mother cells divide is mistaken, as are some of his interpretations. Harris considers other candidates, especially Wilhelm Hofmeister, whose publications in 1848-9 include plates which definitely show mitotic events. Hofmeister was also the choice of Cyril Darlington.
http://en.wikipedia.org/wiki/Chromosomes

It was only in1956 when Joe-Hin Tjio and Albert Levan in Lund, Sweden showed that the correct human chromosome number is 46. http://www.ichg2006.com/abstract/1033.ht…
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